I think that the final antagonists are either going to be Guren and Mahiru, Rígr Stafford or Siga Madu.. The story has made a lot of references to Christian imagery and motifs such as Seraphs, The Four Horsemen of the Apocalypse and Siga Madu alluding to a connection with Lucifer. On the other hand, Ashera Tepes' backstory shows that he lived somewhere in Pre-Christian Europe at around the same period of the Greek Dark Ages- between circa 1100 BC and circa 800 BC....If we assume that this story follows the Judeo-Christian creation story, then I guess that he's probably Lucifer. I did think he could be Prometheus, but Lucifer seems more likely.
Siga Madu says to Rígr Stafford (Saitou), "I have these wings despite losing the sun forever and falling into the darkness, all because of a simple taboo." Is he supposed to be Lucifer? Also, What the actual hell is wrong with Guren and his penchant for being a backstabbing bitch? ヽ(`Д´)ノ