From what I could tell it seemed like the uncle was into chuono’s mom (not sure if they actually had a physical relationship or not), so when she passed away he grieved her and took it too far by basically imposing his affection for her onto her son. In the uncles mind he probably looked similar enough to his mother when he was younger. He basically raped him a bunch and probably left once he got older.
Hey! So there’s actually a short story branching off about chouno’s backstory that literally answers all of your questions!
https://www.mangago.me/read-manga/chou_to_kumo_no_su/

Omg they were so cute the delivery couple was just perfect.
Tw -
I just need someone to answer my question lol about the uncle x the seme with the tattoo. Like what happened, i understood that he essentially groomed by him. However i didn’t really get it that much, i just need more clarification about it sadly ( ̄∇ ̄")